The other players released were Kyle Eastmond, Ben Foden, Alex Goode, Tom Johnson, Freddie Burns and Matt Kvesic, whose absence confirms that Chris Robshaw will play No7.
Scotland could give debuts to London Irish back-row pair Kieran Low and Blair Cowan in their matches against Japan, South Africa and Australia.
Wales’ regions have expressed their desire to play in the rebel Anglo-French Rugby Champions Cup next season instead of the Heineken Cup, subject to WRU approval.
London Irish winger Marland Yarde puts a spring in England"s step
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